I’m glad that you got it working. However,
This should not have been required and you may have found a bug. The intent of Local network on the pjsip tab is to define a per-interface local network, in addition to those already defined in General SIP Settings. Possibly, the pjsip driver is only examining the first General entry.
However, when any of these are changed, one must restart (not just reload) Asterisk for them to take effect.
If you are sure that the necessary restarts were done before you added the pjsip-specific entry, please file a report at issues.freepbx.org .
If not sure, please test whether removing the pjsip-specific Local network (and restarting Asterisk) causes the audio path to stop working.
(The above all assumes that the PBX has just one NIC. If you have multiple, please describe the setup.)